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What is a species that plays an especially important role in it's community called?

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keystone

Some predator species are known as keystone species. A keystone species is one that plays an especially important role in its community. Major changes in the numbers of a keystone species affect the populations of many other species in the community. For example, some sea star species are keystone species in coral reef communities. The sea stars prey on mussels and sea urchins, which have no other natural predators. 

If sea stars were removed from a coral reef community, mussel and sea urchin populations would have explosive growth. This, in turn, would drive out most other species. In the end, the coral reef community would be destroyed.

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