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What are the ultimate source of genetic variation?

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mutations

Mutations and evolution There are often multiple alleles of a particular gene in the population and they all may be equally normal, that is have similar effects on reproductive success andin terms of the phenotypes they produce. If there is no significant selective advantage between them, their relative frequencies within a population will drift. At the same time, the phenotype associated with a particular allelecan be influenced by which alleles are present at other genetic loci, known as the genetic background. Since most traits are the results of hundreds or thousands of genes functioning together, and different combinations of alleles can produce different effects, the universe of variation is large. This can make identifying the genetic basis of a disease difficult, particularly when variation at any one locus may make only a minor contribution to the disease phenotype. On top of that, environmental and developmental differences can outweigh genetic influence on phenotype. Such genetic background effects can lead to a particular allele producing a disease in one person and not another.270 Mutations are the ultimate source of genetic variation  without them evolution would not occur. Mutations can lead to a number of effects, in particular, they can create new activities. At the same time these changes may reduce the original  and necessary  activity of an important gene. Left unresolved such molecular level conflicts would greatly limit the flexibility of evolutionary mechanisms. For example, it is common to think of a gene  or rather the particular gene product it encodes  as having one and only one function or activity, but in fact, when examined closely many catalytic gene products  typically proteins  can catalyze off-target reactions or carry out, even if rather inefficiently, other activities - they interact with other molecules within the cell and the organism. Assume for the moment that a gene encodes a gene product with an essential function as well as potentially useful  from a reproductive success perspective  activities. Mutations that enhance these ancillary functions will survive  that is be passed on to subsequent generations  only to the extent that they do not  overly  negatively influence the genes primary and essential function. The evolution of ancillary functions may be severely constrained or blocked altogether. This problem can be circumvented based on the fact that the genome is not static   . There are molecular level processes through 270.

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